Urea-induced unfolding of Glossoscolex paulistus hemoglobin, in oxy- and cyanomet-forms: A dissociation model

The urea effect on the giant extracellular hemoglobin of Glossoscolex paulistus (HbGp) stability was studied by analytical ultracentrifugation (AUC) and small angle X-ray scattering (SAXS). AUC data show that the sedimentation coefficient distributions curves c (S), at 1.0mol/L of urea, display a si...

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Detalles Bibliográficos
Autores: Carvalho, Francisco A.O., Carvalho, José Wilson P., Santiago, Patrícia S. [UNESP], Tabak, Marcel
Tipo de recurso: artículo
Estado:Versión publicada
Fecha de publicación:2013
País:Brasil
Institución:Universidade Estadual Paulista (UNESP)
Repositorio:Repositório Institucional da UNESP
Idioma:inglés
OAI Identifier:oai:repositorio.unesp.br:11449/74173
Acceso en línea:http://dx.doi.org/10.1016/j.ijbiomac.2012.09.023
http://hdl.handle.net/11449/74173
Access Level:acceso abierto
Palabra clave:AUC
Extracellular hemoglobin
HbGp
Oligomeric dissociation
SAXS
Urea
cyanomethemoglobin
hemoglobin
oxyhemoglobin
unclassified drug
urea
annelid
concentration response
dissociation
Glossoscolex paulistus
molecular weight
nonhuman
oligomeric dissociation
protein stability
protein structure
protein unfolding
sedimentation
ultracentrifugation
X ray crystallography
Animals
Hemoglobins
Models, Chemical
Oligochaeta
Protein Folding
Glossoscolex
Descripción
Sumario:The urea effect on the giant extracellular hemoglobin of Glossoscolex paulistus (HbGp) stability was studied by analytical ultracentrifugation (AUC) and small angle X-ray scattering (SAXS). AUC data show that the sedimentation coefficient distributions curves c (S), at 1.0mol/L of urea, display a single peak at 57 S, associated to the undissociated protein. The increase in urea concentration, up to 4.0mol/L, induces the appearance of smaller species, due to oligomeric dissociation. The sedimentation coefficients and molecular masses are 9.2S and 204kDa for the dodecamer (abcd)3, 5.5S and 69kDa for the tetramer (abcd), 4.1S and 52kDa for the trimer (abc) and 2.0 S and 17kDa for the monomer d, respectively. SAXS data show initially a decrease in the I(0) values due to the oligomeric dissociation, and then, above 4.0mol/L of denaturant, for oxy-HbGp, and above 6.0mol/L for cyanomet-HbGp, an increase in the maximum dimension and gyration radius is observed, due to the unfolding process. According to AUC and SAXS data the HbGp unfolding is described by two phases: the first one, at low urea concentration, below 4.0mol/L, characterizes the oligomeric dissociation, while the second one, at higher urea concentration, is associated to the unfolding of dissociated species. Our results are complementary to a recent report based on spectroscopic observations. © 2012 Elsevier B.V.